NK cell and cancer cell




Pradeu T. (2012), The Limits of the Self: Immunology and Biological Identity. New York: Oxford University Press. Lakatos Award 2015. Soon available in paperback (May 2018).

(Oxford Scholarship Online).

Reviews in: Notre Dame Philosophical Reviews (Alfred I. Tauber); Times Literary Supplement (R. P. Novick); Studies in History and Philosophy of Biological and Biomedical Sciences (Neeraja Sankaran); British Journal for the Philosophy of Science (Maureen O'Malley); Biology and Philosophy (Matt Haber); Philosophy in Review (Alison McConwell).

Carosella E.D., Pradeu T. (2010), L'Identité: La Part de l'Autre. Paris: Odile Jacob.

Pradeu T. (2009), Les Limites du soi: Immunologie et identité biologique. Montréal: Presses Universitaires de Montréal, & Paris: Vrin. Compte rendu dans History and Philosophy of the Life Sciences (F. Bouchard)



Edited volumes

Gayon J., and Pradeu T. (eds.) (in press), Textes Clés de Philosophie de la Biologie (Key readings in Philosophy of Biology), 2 vol., Paris,  Vrin.

Guay A. and Pradeu T. (eds.) (2016) Individuals Across the Sciences. New York: Oxford University Press. Contributors: Stéphane Chauvier; Ruey-Lin Chen; Christina Conroy; Mark Ereshefsky & Makmiller Pedroso; Melinda Fagan; Steven French; David Glick; Peter Godfrey-Smith; Alexandre Guay & Thomas Pradeu; Matt Haber; Décio Krause & Jonas R. Becker Arenhart; James Ladyman; Jonathan E. Lowe; Matteo Morganti; Cédric Paternotte; Paavo Pylkkanen, Basil J. Hiley & Ilkka Pattiniemi; Simon Saunders.

Individuals Across the Sciences_Cover Design

Minelli A., Pradeu T., eds. (2014) Towards a Theory of Development. Oxford: Oxford University Press. Contributors: Wallace Arthur; Giuseppe Fusco, Roberto Carrer and Emanuele Serrelli; Scott Gilbert and Jonathan Bard; James Griesemer; Isaac Hernández and Davide Vecchi; Johannes Jaeger and James Sharpe; Jean-Jacques Kupiec; Lucie Laplane; Alan Love; Alessandro Minelli; Armin P. Moczek; Michel Morange; Stuart Newman; Spencer V. Nyholm and Margaret McFall-Ngai; Thomas Pradeu; Michel Vervoort.
minelli&pradeu_Theory dvpt
Barberousse A., Morange M., Pradeu T., eds. (2009) Mapping the Future of Biology: Evolving Concepts and Theories. Dordrecht: Springer, Boston Studies in the Philosophy and History of Science, vol. 266. (Contributors: Mark A. Bedau, Evelyn Fox Keller, Scott F. Gilbert, Peter Godfrey-Smith, Michel Morange, John Odling-Smee, Susan Oyama, Michael Ruse, and Kim Sterelny)
Ludwig P., Pradeu T., eds. (2008), L'Individu: Perspectives contemporaines. Paris: Vrin. Spanish translation, El individuo. Perspectivas contemporáneas, Nueva Vision, 2014.
Carosella E.D., Pradeu T., Saint-Sernin B., Debru C., eds. (2006) L'identité? Soi et non-soi, individu et personne. Paris, Presses Universitaires de France.



Edition of special issues




  • Pradeu T., Kostyrka G., and Dupré J. (eds., 2016), Special issue of Studies C entitled "Understanding Viruses: Philosophical Investigations", Studies in History and Philosophy of Science Part C: Studies in History and Philosophy of Biological and Biomedical Sciences 59 (October 2016), 57-153. Contributors: Jean-Michel Claverie & Chantal Abergel, John Dupré, Patrick Forterre, Eugene Koonin & Petro Starokadomskyy, Gladys Kostyrka, Pierre-Olivier Méthot, Greg Morgan, Maureen O’Malley, Thomas Pradeu, and Marc van Regenmortel.

Understanding viruses_Front pageVirus_sn-hepatitis


  • Guest editor, Biological Theory: issue on "The Boundaries of development" (6:1, Dec 2011), with the contributions of Scott Gilbert, Lucie Laplane, Jane Maienschein, Alessandro Minelli, Michel Morange, Antonine Nicoglou, Thomas Pradeu, Frédérique Théry, and Michel Vervoort.



Mutualistic viruses_Front pageParasitoid wasp


  • Pradeu T., Kostyrka G., and Dupré J (2016), Understanding viruses: Philosophical investigations, Studies in History and Philosophy of Biological and Biomedical Sciences 59, 57-63. (PDF of final draft)

  • Pradeu T, Laplane L, Prévot K, Hoquet T, Reynaud V, Fusco G, Minelli A, Orgogozo V, and Vervoort M. (2016), Defining 'development', Current Topics in Developmental Biology, 117, 171-183, special anniversary issue edited by Paul Wassarman. (PDF of final draft)

Pradeu-Jaeger-Vivier_Speed of change_Front page

'Monitoring discontinuities' by Simon Bradbrook, inspired by the Review on p764.







Articles in French


Book chapters

Pradeu T. (in press), Philosophy of Biology, in. A. Barberousse, D. Bonnay & M. Cozic (eds.), The Philosophy of Science. Oxford University Press.

Pradeu T. (2018), Jean Gayon, l’histoire, la philosophie et la biologie : Une synthèse moderne, in F. Merlin & Ph. Huneman (eds.), Philosophie, histoire, biologie. Mélanges offerts à Jean Gayon. Paris, Editions Matériologiques.

Pradeu T. (2018), Genidentity and Biological Processes, in Nicholson, D. J. & Dupré, J. (eds.). Everything Flows: Towards a Processual Philosophy of Biology. Oxford: Oxford University Press. Open access to the whole book in PDF file.

Guay A. and Pradeu T. (2016), Introduction: Progressive Steps Towards a Unified Conception of Individuality Across the Sciences, in Guay and Pradeu, eds., Individuals Across the Sciences, pp. 1-21.

Guay A. and Pradeu T. (2016), To be continued: the genidentity of physical and biological processes, in Guay and Pradeu, eds., Individuals Across the Sciences, pp. 317-347.

Minelli A. & Pradeu T. (2014), Theories of development in biology – problems and perspectives, in A. Minelli and T. Pradeu, eds., Towards a Theory of Development. Oxford: Oxford University Press, pp. 1-14.

Pradeu T. (2014), Regenerating Theories in Developmental Biology, in A. Minelli and T. Pradeu (eds.) Towards a Theory of Development. Oxford: Oxford University Press, pp. 15-32.

Pradeu T. (2013), Immunity and the emergence of individuality, in F. Bouchard and P. Huneman (eds.) From Groups to Individuals: Perspectives on Biological Associations and Emerging Individuality. Cambridge, MA: MIT Press, pp. 77-96.


Pradeu T. (2013), Critique des notions de soi et de non-soi en immunologie, in J-J. Kupiec (ed.) La Vie, et alors? Paris: Belin, pp. 177-189.

Kupiec_Cover_La vie et alors

Pradeu T. (2011), Chapitre "Philosophie de la biologie", in A. Barberousse, D. Bonnay, M. Cozic (eds.) Précis de philosophie des sciences. Paris, Vuibert, 378-403.


Pradeu T. (2010), Peut-on attribuer une fonction au système immunitaire?, in J. Gayon & A. de Ricqlès (eds.) Les Fonctions : des organismes aux artefacts. Paris: Presses Universitaires de France, pp. 261-275.

Couv_Fonctions des organismes aux artefacts

Pradeu T. (2009), Darwinisme, évolution et immunologie, in T. Heamns, P. Huneman, G. Lecointre, M. Silberstein (eds.), Les Mondes darwiniens: L'évolution de l'évolution. Paris: Syllepses, pp. 525-542.

Site web:

Nouvelle édition parue en 2011 aux éditions Matériologiques: 


Pradeu T. (2008), Qu’est-ce qu’un individu biologique?, in P. Ludwig, T. Pradeu (eds.), L’individu : Perspectives contemporaines. Paris : Vrin, pp. 97-125.

Pradeu T. (2007), L'immunité et l'interactionnisme biologique, in Thierry Martin (dir.), Le Tout et les parties dans les systèmes naturels. Paris: Vuibert, coll. "Philosophie des sciences", pp. 99-106.

Tout et parties

Carosella E.D., Pradeu T. (2006), Introduction, in Carosella E.D., Pradeu T., Saint-Sernin B., Debru C. (eds.) L'identité? Soi, non-soi, individu et personne. Paris: PUF, pp. 3-5.

Pradeu T., Carosella E.D. (2006), L'identité immunologique : soi ou continuité?, in Carosella E.D., Pradeu T., Saint-Sernin B., Debru C. (eds.) L'identité? Soi, non-soi, individu et personne. Paris: PUF, pp. 47-58.




Book reviews



Outreach activities





Work in progress

  • Pradeu T. & Moreau J-F. (submitted), CRISPR-Cas immunity: Beyond nonself and defence. Submitted to Biology & Philosophy.
  • Pradeu T. (in prep.), Philosophy of CRISPR-Cas systems. In preparation for Biology & Philosophy.
  • Chiu L. & Pradeu T. (submitted) Extension Arguments across the Biological Sciences.
  • Pradeu T. Philosophy of Immunology. Cambridge University Press.
  • Pradeu T. & Ferner A. (in prep.), A Critique of animalism.